Origin of Species and Macroevolution
Key Concepts:
Identification of Species
Mechanisms of Speciation
The Pace of Speciation
Evo-Devo: Evolutionary Developmental Biology
Identification of Species
Macroevolution
Evolutionary changes that create new species and groups of species
Concerns about the diversity of organisms established over long periods of time through the evolution and extinction of many species
Species
A group of organisms that maintains a distinctive set of attributes in nature
Currently, about 1.75 million species identified
Estimates of the total number of species range from 5 - 50 million
Difficulty in identifying a “species”
Subspecies
Ecotypes
Amount of separation time for two populations
Short time – likely to be similar and considered the same species
Long time – more likely to show explicit differences
May find situations where some differences are apparent but difficult to decide if the two populations are truly different species
Sometimes use subspecies classification
Characteristics that a biologist uses to identify a species will depend, in large part, on the species in question
Most commonly used characteristics are
morphological traits,
ability to interbreed,
molecular features,
ecological factors, and
evolutionary relationships
Physical characteristics of an organism
Drawbacks for determining species
How many traits to consider
Traits may vary in a continuous way; quantitative
What degree of dissimilarity to use
Members of the same species can look very different
Members of a different species can look very similar
Reproductive isolation
Prevents one species from successfully interbreeding with other species
Four main problems for determining species
•May be difficult to determine in nature
•Can interbreed and yet do not
•Does not apply to asexual species
•Cannot be applied to extinct species